BrainTrain project publications related to learning, memory, plasticity and/or executive functioning

In press

  • Fellman, D., Salmi, J., Ritakallio, L., Ellfolk, U., Rinne, J., & Laine, M. (in press). Training working memory updating in Parkinson’s disease: A randomized controlled trial. Neuropsychological Rehabilitation. doi:10.1080/09602011.2018.1489860. [Epub ahead of print]
  • Fellman, D., Jylkkä, J., Waris, O., Soveri, A., Ritakallio, L., Haga, S., Salmi, J., Nyman, T.N., & Laine, M. (in press). The role of strategy use in working memory training outcomes. Journal of Memory and Language. doi:10.1016/j.jml.2019.10406
  • Salmi, J., Metwalye, M., Tohka, J., Alho, K., Leppämäki, S., Tani, P., Koski, A., Vanderwal, T., & Laine, M. (in press). ADHD desynchronizes brain activity during watching a distracted multi-talker conversation. NeuroImage. doi:10.1016/j.neuroimage.2019.116352
  • Salmi, J., Vilà-Balló, A., Soveri, A., Rostan, C., Rodríguez-Fornells, A., Lehtonen, M., & Laine, M. (in press). Working memory updating training modulates a cascade of event-related potentials depending on task load. Neurobiology of Learning and Memory, 9, 107085. doi:10.1016/j.nlm.2019.107085. [Epub ahead of print]


  • Lukasik, K.M., Waris, O., Soveri, A., Lehtonen, M., & Laine, M. (2019) The relationship of anxiety and stress with working memory performance in a large non-depressed sample. Frontiers in Psychology, 10(4). doi:10.3389/fpsyg.2019.00004
  • Waris, O., Jaeggi, S.M., Seitz, A.R., Lehtonen, M., Soveri, A., Lukasik, K.M., Söderström, U., Hoffing, R.A.C., & Laine, M. (2019) Video gaming and working memory: A large-scale cross-sectional correlative study. Computers in Human Behavior, 97, 94-103. doi:10.1016/j.chb.2019.03.005


  • Fellman, D., Soveri, A., Viktorsson, C., Haga, S., Nylund, J., Johansson, S., Edman, J., von Renteln, F., & Laine, M. (2018) Selective updating of sentences: Introducing a new measure of verbal working memory. Applied Psycholinguistics, 39, 275-301. doi:10.1017/S0142716417000182
  • Jylkkä, J., Lehtonen, M., Kuusakoski, A., Lindholm, F., Hut, S., & Laine, M. (2018). The role of general executive functions in receptive language switching and monitoring. Bilingualism: Language and Cognition, 21, 839-855. doi:10.1017/S1366728917000384
  • Laine, M., Fellman, D., Waris, O., & Nyman T.N. (2018) The early effects of external and internal strategies on working memory updating training. Scientific Reports, 8, 4045. doi:10.1038/s41598-018-22396-5
  • Lehtonen, M., Soveri, A., Laine, A., Järvenpää, J., De Bruin, A. & Antfolk, J. (2018). Is bilingualism associated with enhanced executive functioning? A meta-analytic review. Psychological Bulletin, 144, 394-425. doi:10.1037/bul0000142
  • Lukasik, K.M., Lehtonen, M., Salmi, J., Meinzer, M., Joutsa, J., & Laine, M. (2018) No effects of stimulating the left ventrolateral prefrontal cortex with tDCS on verbal working memory updating. Frontiers in Neuroscience, 1, 738. doi:10.3389/fnins.2017.00738
  • Lukasik, K.M., Lehtonen, M., Soveri, A., Waris, O., Jylkkä, J. & Laine, M. (2018) Bilingualism and working memory performance: evidence from a large-scale online study. PLoS ONE, 13. doi:10.1371/journal.pone.0205916.
  • Salmi J., Nyberg, L., & Laine, M. (2018) Working memory training mostly engages general-purpose large-scale networks for learning. Neuroscience and Biobehavioral Reviews, 93, 108-122. doi:10.1016/j.neubiorev.2018.03.019
  • Soveri, A., Lehtonen, M., Karlsson, L.C., Lukasik, K., Antfolk, A., & Laine, M. (2018) Test-retest reliability of five frequently used executive tasks in healthy adults. Applied Neuropsychology: Adult, 25, 155-165. doi:10.1080/23279095.2016.1263795
  • Vilà-Balló, A., Salmi, J., Soveri, A., Rodríguez-Fornells, A., Lehtonen, M., & Laine, M. (2018) Neural signatures for active maintenance and interference during working memory updating. Biological Psychology, 132, 233-243. doi:10.1016/j.biopsycho.2018.01.007
  • Waris, O., Soveri, A., Lukasik, K.M., Lehtonen, M., & Laine, M. (2018) Working memory and the Big Five. Personality and Individual Differences, 130, 26-35. doi:10.1016/j.paid.2018.03.027


  • Bäckman, L., Waris, O., Johansson, J., Andersson, M., Rinne, J.O., Alakurtti, K., Soveri, A., Laine, M., & Nyberg, L. (2017) Increased dopamine release after working-memory updating training: Neurochemical correlates of transfer. Scientific Reports, 7. doi:10.1038/s41598-017-07577-y
  • Fellman, D., Soveri, A., Waris, O., & Laine, M. (2017) Training of verbal working memory at sentence level fails to show transfer. Frontiers in Communication, 2(14). doi:10.3389/fcomm.2017.00014.
  • François, C., Cunillera, T., Garcia, E., Laine, M., & Rodriguez-Fornells, A. (2017) Neurophysiological evidence for the interplay of speech segmentation and word-referent mapping during novel word learning. Neuropsychologia, 98, 56-67. doi:10.1016/j.neuropsychologia.2016.10.006
  • Grönholm-Nyman, P., Soveri, A., Rinne, J.O., Ek, E., Nyholm, A., Stigsdotter Neely, A., & Laine, M. (2017) Limited effects of set shifting training in healthy older adults. Frontiers in Aging Neuroscience, 9(69), 1-21. doi:10.3389/fnagi.2017.00069
  • Havas, V., Laine, M., & Rodríguez Fornells, A. (2017) Brain signatures of early lexical and morphological learning of a new language. Neuropsychologia, 101, 47-56. doi:10.1016/j.neuropsychologia.2017.04.005
  • Jylkkä, J., Soveri, A., Wahlström, J., Lehtonen, M., Rodríguez-Fornells, A., & Laine, M. (2017) Relationship between language switching experience and executive functions in bilinguals: an Internet-based study. Journal of Cognitive Psychology, 29, 404-419. doi:10.1080/20445911.2017.1282489
  • Saarela, C., Joutsa, J., Laine, M., Parkkola, R., Rinne, J.O., & Karrasch, M. (2017) Regional gray matter correlates of memory for emotion-laden words in middle-aged and older adults: A voxel-based morphometry study. PLoS ONE, 12(8), e0182541. doi:10.1371/journal.pone.0182541
  • Soveri, A., Antfolk, J., Karlsson, L.C., Salo, B., & Laine, M. (2017) Working memory training revisited: a multi-level meta-analysis of n-back training studies. Psychonomic Bulletin & Review, 24, 1077–1096. doi:10.3758/s13423-016-1217-0
  • Soveri, A., Karlsson, E.P.A., Waris, O., Grönholm-Nyman, P., & Laine, M. (2017) Pattern of near transfer effects following working memory training with a dual n-back task. Experimental Psychology, 64, 240–252. doi:10.1027/1618-3169/a000370
  • Tort-Merino, A., Valech, N., Peñaloza, C., Grönholm-Nyman, P., León, M., Olives, J., Estanga, A., Ecay, M., Fortea, J., Martínez-Lage, P., Molinuevo, J. L., Laine, M., Rodríguez-Fornells, A., & Rami, L. (2017) Early detection of learning difficulties when confronted with novel information in preclinical Alzheimer's Disease stage 1. Journal of Alzheimer’s Disease, 58, 855–870. doi:10.3233/JAD-161173
  • Waris, O., Soveri, A., Ahti, M., Hoffing, R.C., Ventus, D., Jaeggi, S.M., Seitz, A.R., & Laine, M. (2017) A latent factor analysis of working memory measures using large-scale data. Frontiers in Psychology, 8. doi:10.3389/fpsyg.2017.01062.


  • Cunillera, T., Laine, M., & Rodríguez-Fornells, A. (2016) Headstart for speech segmentation: a neural signature for the anchor word effect. Neuropsychologia, 82, 189-199. doi:10.1016/j.neuropsychologia.2016.01.011
  • Peñaloza, C., Mirman, D., Tuomiranta, L., Benetello, A., Heikius, I.-M., Järvinen, S., Majos, M.C., Cardona, P., Juncadella, M., Laine, M., Martin, N., & Rodríguez-Fornells, A. (2016) Novel word acquisition in aphasia: facing the word-referent ambiguity of natural language learning contexts. Cortex, 79, 14-31. doi:10.1016/j.cortex.2016.03.009
  • Saarela, C., Karrasch, M., Ilvesmäki, T., Parkkola, R., Rinne, J.O., & Laine, M. (2016) The relationship between recognition memory for emotion-laden words and white matter microstructure in normal older subjects. Neuroreport, 27, 1345-1347. doi:10.1097/WNR.0000000000000704


  • Havas, V., Waris, O., Vaquero, L., Rodríguez-Fornells, A., & Laine, M. (2015) Morphological learning in a novel language: a cross-language comparison. Quarterly Journal of Experimental Psychology, 68, 1426-1441. doi:10.1080/17470218.2014.983531
  • Karlsson, L.C., Soveri, A., Räsänen, P., Kärnä, A., Delatte, S., Lagerström, E., Mård, L., Steffansson, M., Lehtonen, M., & Laine, M. (2015) Bilingualism and performance on two widely used developmental neuropsychological test batteries. PLoS ONE, 10(4), e0125867, 1-11. doi:10.1371/journal.pone.0125867
  • Martin, N., Laine, M. & Harley, T.A. (2015) How can connectionist cognitive models of language inform models of language rehabilitation? In A.E. Hillis (Ed.) Handbook on Adult Language Disorders. Second Edition. New York: Psychology Press (pp. 405-434).
  • Peñaloza, C., Benetello, A., Tuomiranta, L., Heikius, I.-M., Järvinen, S., Majos, M.C., Cardona, P., Juncadella, M., Laine, M., Martin, N., & Rodríguez-Fornells, A. (2015) Speech segmentation in aphasia. Aphasiology, 29, 724-743. doi:10.1080/02687038.2014.982500
  • Tallus, J., Soveri, A., Hämäläinen, H., Tuomainen, J., & Laine, M. (2015) Effects of auditory attention training with the dichotic listening task: Behavioural and neurophysiological evidence. PLoS ONE, 1-14. doi:10.1371/journal.pone.0139318
  • Waris, O., Soveri, A., & Laine, M. (2015) Transfer after working memory updating training. PLoS ONE, 1-19. doi:10.1371/journal.pone.0138734


Department of Psychology
Abo Akademi University
Fabriksgatan 2
FI-20500 Turku